Rodriguez, L. O., & Catenazzi, A. (2017).

Four new species of terrestrial-breeding frogs of the genus Phrynopus (Anura: Terrarana: Craugastoridae) from Río Abiseo National Park, Peru

. Zootaxa, 4273(3), 381–406. https://doi.org/10.11646/zootaxa.4273.3.4