Kluge, N. J., Godunko, R. J., & Apanaskevich, D. A. (2013). Mayflies of the Caucasus Mountains. II. Description of the first representative of the subgenus Helvetoraeticus Bauernfeind & Soldán, 2012 (Heptageniidae: Ecdyonurus). Zootaxa, 3608(1), 51–66. https://doi.org/10.11646/zootaxa.3608.1.3