Maddison, D. R., Toledano, L., Sallenave, S., & Roig-Juñent, S. (2013). Phylogenetic relationships of the South American ground beetle subgenus Chilioperyphus Jeannel (Coleoptera: Carabidae: Trechinae: Bembidiini: Bembidion Latreille). Zootaxa, 3636(4), 547–560. https://doi.org/10.11646/zootaxa.3636.4.3