Angulo, A., & Gracian-Negrete, J. M. (2013). A new species of Brycon (Characiformes: Characidae) from Nicaragua and Costa Rica, with a key to the lower Mesoamerican species of the genus. Zootaxa, 3731(2), 255–266. https://doi.org/10.11646/zootaxa.3731.2.6