Bernal, M. H., Luna-Mora, V. F., Gallego, O., & Quevedo, A. (2007).

A new species of poison frog (Amphibia: Dendrobatidae) from the Andean mountains of Tolima, Colombia

. Zootaxa, 1638(1), 59–68. https://doi.org/10.11646/zootaxa.1638.1.5