Lobo, J. M., Jiménez-Ruiz, Y., Chehlarov, E., Guéorguiev, B., Petrova, Y., Král, D., Alonso-Zarazaga, M. Ángel, & Verdú, J. R. (2015). The classification and phylogenetic status of Jekelius (Reitterius) punctulatus (Jekel, 1866) and Jekelius (Jekelius) brullei (Jekel, 1866) (Coleoptera: Geotrupidae) using molecular data. Zootaxa, 4040(2), 187–203. https://doi.org/10.11646/zootaxa.4040.2.5