Twomey, E., & Brown, J. L. (2008). A partial revision of the Ameerega hahneli complex (Anura: Dendrobatidae) and a new cryptic species from the East-Andean versant of Central Peru. Zootaxa, 1757(1), 49–65. https://doi.org/10.11646/zootaxa.1757.1.3