García-López, A., Morón, M. Ángel, Micó, E., & Galante, E. (2009). Two new species of Phyllophaga Harris (Coleoptera: Scarabaeidae: Melolonthinae) from Costa Rica. Zootaxa, 2062(1), 37–45. https://doi.org/10.11646/zootaxa.2062.1.3