Matsumoto, T., Matsuura, K., & Hanzawa, N. (2021).

A new species of nine-spined stickleback, Pungitius modestus (Gasterosteiformes, Gasterosteidae), from northern Honshu, Japan

. Zootaxa, 5005(1), 1–20. https://doi.org/10.11646/zootaxa.5005.1.1