GASCA-ÁLVAREZ, H. J.; RATCLIFFE, B. C. An annotated checklist of the oryctine rhinoceros beetles (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) of the Neotropical and Nearctic realms. Zootaxa, Auckland, New Zealand, v. 3090, n. 1, p. 21–40, 2011. DOI: 10.11646/zootaxa.3090.1.2. Disponível em: https://mapress.com/zt/article/view/zootaxa.3090.1.2. Acesso em: 9 jun. 2025.