SOLÍS, Ángel; KOHLMANN, B. Checklist and distribution atlas of the Scarabaeinae (Coleoptera: Scarabaeidae) of Costa Rica. Zootaxa, Auckland, New Zealand, v. 3482, n. 1, p. 1–32, 2012. DOI: 10.11646/zootaxa.3482.1.1. Disponível em: https://mapress.com/zt/article/view/zootaxa.3482.1.1. Acesso em: 20 apr. 2025.