NUNES, R. V.; VAZ-DE-MELLO, F. Z. A new species of Holocephalus Hope from Paraguay (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini s. str.). Zootaxa, Auckland, New Zealand, v. 4136, n. 2, p. 397–400, 2016. DOI: 10.11646/zootaxa.4136.2.12. Disponível em: https://mapress.com/zt/article/view/zootaxa.4136.2.12. Acesso em: 22 apr. 2025.