LÓPEZ-GARCÍA, M. M.; GASCA-ÁLVAREZ, H. J.; CAVE, R. D.; AMAT-GARCÍA, G. An annotated checklist of the New World pentodontine scarab beetles (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini) . Zootaxa, Auckland, New Zealand, v. 4170, n. 3, p. 491–509, 2016. DOI: 10.11646/zootaxa.4170.3.4. Disponível em: https://mapress.com/zt/article/view/zootaxa.4170.3.4. Acesso em: 22 apr. 2025.