KOHLMANN, B.; SOLÍS, Ángel. New species of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) from Mexico and Costa Rica. Zootaxa, Auckland, New Zealand, v. 1302, n. 1, p. 61–68, 2006. DOI: 10.11646/zootaxa.1302.1.5. Disponível em: https://mapress.com/zt/article/view/zootaxa.1302.1.5. Acesso em: 19 apr. 2025.