RATCLIFFE, B. C. A checklist of the Scarabaeoidea (Coleoptera) of Panama. Zootaxa, Auckland, New Zealand, v. 32, n. 1, p. 1–48, 2002. DOI: 10.11646/zootaxa.32.1.1. Disponível em: https://mapress.com/zt/article/view/zootaxa.32.1.1. Acesso em: 4 apr. 2025.