MEDINA, C. A.; MOLANO, F.; SCHOLTZ, C. H. Morphology and terminology of dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) male genitalia. Zootaxa, Auckland, New Zealand, v. 3626, n. 4, p. 455–476, 2013. DOI: 10.11646/zootaxa.3626.4.3. Disponível em: https://mapress.com/zt/article/view/zootaxa.3626.4.3. Acesso em: 21 apr. 2025.