CUPELLO, M.; VAZ-DE-MELLO, F. Z. Taxonomic revision of the South American dung beetle genus Gromphas Brullé, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina). Zootaxa, Auckland, New Zealand, v. 3722, n. 4, p. 439–482, 2013. DOI: 10.11646/zootaxa.3722.4.2. Disponível em: https://mapress.com/zt/article/view/zootaxa.3722.4.2. Acesso em: 24 apr. 2025.