LÓPEZ-GARCÍA, M. M.; GASCA-ÁLVAREZ, H. J.; AMAT-GARCÍA, G. A new species of Tomarus Erichson, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini), with a key to the species in Colombia. Zootaxa, Auckland, New Zealand, v. 3869, n. 5, p. 579–584, 2014. DOI: 10.11646/zootaxa.3869.5.8. Disponível em: https://mapress.com/zt/article/view/zootaxa.3869.5.8. Acesso em: 30 apr. 2025.