LÓPEZ-GARCÍA, M. M.; GASCA-ÁLVAREZ, H. J.; AMAT-GARCÍA, G.

The scarab beetle tribe Pentodontini (Coleoptera: Scarabaeidae: Dynastinae) of Colombia: taxonomy, natural history, and distribution

. Zootaxa, Auckland, New Zealand, v. 4048, n. 4, p. 451–492, 2015. DOI: 10.11646/zootaxa.4048.4.1. Disponível em: https://mapress.com/zt/article/view/zootaxa.4048.4.1. Acesso em: 22 apr. 2025.