SHAPOVALOV, A. M.; BAGATUROV, M. F. A review of the subgenus Mesolethrus Nikolajev, 2003 of the genus Lethrus Scopoli, 1777 (Coleoptera: Geotrupidae: Lethrinae) with description of a new species. Zootaxa, Auckland, New Zealand, v. 5351, n. 4, p. 435–452, 2023. DOI: 10.11646/zootaxa.5351.4.2. Disponível em: https://mapress.com/zt/article/view/zootaxa.5351.4.2. Acesso em: 4 apr. 2025.