GUAYASAMIN, J. M.; BONACCORSO, E.; DUELLMAN, W. E.; COLOMA, L. A. Genetic differentiation in the nearly extinct harlequin frogs (Bufonidae: Atelopus), with emphasis on the Andean Atelopus ignescens and A. bomolochos species complexes. Zootaxa, Auckland, New Zealand, v. 2574, n. 1, p. 55–68, 2010. DOI: 10.11646/zootaxa.2574.1.2. Disponível em: https://mapress.com/zt/article/view/zootaxa.2574.1.2. Acesso em: 3 apr. 2025.