Bravo-Valencia, Laura, and Mauricio Rivera-Correa. 2011. “A New Species of Harlequin Frog (Bufonidae: Atelopus) With an Unusual Behavior from Andes of Colombia”. Zootaxa 3045 (1). Auckland, New Zealand:57–67. https://doi.org/10.11646/zootaxa.3045.1.4.