Brown, Jason L., Evan Twomey, Adolfo Amézquita, Moisés Barbosa de Souza, Janalee P. Caldwell, Stefan Lötters, Rudolf Von May, Paulo Roberto Melo-Sampaio, Daniel Mejía-Vargas, Pedro Perez-Peña, Mark Pepper, Erik H. Poelman, Manuel Sanchez-Rodriguez, and Kyle Summers. 2011. “A Taxonomic Revision of the Neotropical Poison Frog Genus Ranitomeya (Amphibia: Dendrobatidae)”. Zootaxa 3083 (1). Auckland, New Zealand:1–120. https://doi.org/10.11646/zootaxa.3083.1.1.