Hutter, Carl R., and Juan M. Guayasamin. 2012. “A New Cryptic Species of Glassfrog (Centrolenidae: Nymphargus) from Reserva Las Gralarias, Ecuador”. Zootaxa 3257 (1). Auckland, New Zealand:1–21. https://doi.org/10.11646/zootaxa.3257.1.1.