Arce-Pérez, Roberto, and Miguel Angel Morón. 2005. “New Species and New Records of Species of Macrodactylus Dejean (Coleoptera: Scarabaeidae: Melolonthinae: Macrodactyini) from Honduras and Nicaragua”. Zootaxa 1012 (1). Auckland, New Zealand:23–37. https://doi.org/10.11646/zootaxa.1012.1.3.