Jansen, Martin, and Arne Schulze. 2012. “Molecular, Morphology and Bioacoustic Data Suggest Bolivian Distribution of a Large Species of the Leptodactylus Pentadactylus Group (Amphibia: Anura: Leptodactylidae)”. Zootaxa 3307 (1). Auckland, New Zealand:35–47. https://doi.org/10.11646/zootaxa.3307.1.2.