Kluge, Nikita J., Roman J. Godunko, and Dmitry A. Apanaskevich. 2013. “Mayflies of the Caucasus Mountains. II. Description of the First Representative of the Subgenus Helvetoraeticus Bauernfeind & Soldán, 2012 (Heptageniidae: Ecdyonurus)”. Zootaxa 3608 (1). Auckland, New Zealand:51–66. https://doi.org/10.11646/zootaxa.3608.1.3.