Valencia, Jorge H., Francisco Valladares-Suntasig, Luis Tipantiza-Tuguminago, and Manuel R. Dueñas. 2019. “A New Species of Terrestrial-Breeding Frog of the Genus Pristimantis (Anura: Terrarana: Craugastoridae) from the Eastern Andean Slopes of the Southern Ecuador”. Zootaxa 4658 (3). Auckland, New Zealand:509–525. https://doi.org/10.11646/zootaxa.4658.3.4.