Amézquita, Adolfo, Roberto Márquez, Ricardo Medina, Daniel Mejía-Vargas, Ted R. Kahn, Gustavo Suárez, and Luis Mazariegos. 2013. “A New Species of Andean Poison Frog, Andinobates (Anura: Dendrobatidae), from the Northwestern Andes of Colombia”. Zootaxa 3620 (1). Auckland, New Zealand:163–178. https://doi.org/10.11646/zootaxa.3620.1.8.