Benine, Ricardo C., and Guilherme A. M. Lopes. 2008. “A New Species of Hyphessobrycon (Characiformes: Characidae) from Rio Portuguesa, Rio Orinoco Basin, Venezuela”. Zootaxa 1747 (1). Auckland, New Zealand:61–68. https://doi.org/10.11646/zootaxa.1747.1.3.