Bolaños, Federico, and David B. Wake. 2009. “Two New Species of Montane Web-Footed Salamanders (Plethodontidae: Bolitoglossa) from the Costa Rica-Panamá Border Region”. Zootaxa 1981 (1). Auckland, New Zealand:57–68. https://doi.org/10.11646/zootaxa.1981.1.5.