Bartels, Paul J., Giovanni Pilato, Oscar Lisi, and Diane R. Nelson. 2009. “Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee/North Carolina, USA (North America): Two New Species and Six New Records”. Zootaxa 2022 (1). Auckland, New Zealand:45–57. https://doi.org/10.11646/zootaxa.2022.1.4.