Bitar, Alberto, and Miguel Ángel Morón. 2009. “Description of the Female of Tehuacania Howdeni Endrödi (Coleoptera: Scarabaeidae: Dynastinae: Oryctini), With a Key to the Genera of Oryctini from Mexico”. Zootaxa 2048 (1). Auckland, New Zealand:65–68. https://doi.org/10.11646/zootaxa.2048.1.4.