Flowers, R. Wills. 2009. “A New Species of Thraulodes (Ephemeroptera: Leptophlebiidae, Atalophlebiinae) from a Highly Altered River in Western Ecuador”. Zootaxa 2052 (1). Auckland, New Zealand:55–61. https://doi.org/10.11646/zootaxa.2052.1.4.