Grossi, Paschoal Coelho. 2009. “Description of Two New Species of Leptinopterus Hope (Coleoptera: Lucanidae: Lucaninae) With Notes on the Taxonomy and Natural History of the Genus”. Zootaxa 2172 (1). Auckland, New Zealand:32–44. https://doi.org/10.11646/zootaxa.2172.1.2.