Erwin, Terry L. 2004. “The Beetle Family Carabidae of Costa Rica and Panam : Descriptions of Four New Genera and Six New Species With Notes on Their Way of Life (Insecta: Coleoptera)”. Zootaxa 537 (1). Auckland, New Zealand:1–18. https://doi.org/10.11646/zootaxa.537.1.1.