Pyron, R. Alexander, and David A. Beamer. 2023. “Systematic Revision of the Spotted and Northern Dusky Salamanders (Plethodontidae: Desmognathus Conanti and D. fuscus), With Six New Species from the Eastern United States”. Zootaxa 5311 (4). Auckland, New Zealand:451-504. https://doi.org/10.11646/zootaxa.5311.4.1.