Deschodt, Christian M., and Catherine L. Sole. 2024. “A New and Unusually Ornate Southern African Dung Beetle Species in d’Orbigny’s Onthophagus Group 18 (Coleoptera: Scarabaeidae: Onthophagini)”. Zootaxa 5493 (2). Auckland, New Zealand:186-94. https://doi.org/10.11646/zootaxa.5493.2.7.