Moreno, Jhon C. Neita, Julián Clavijo-Bustos, and Brett C. Ratcliffe. 2025. “A Review of the Genus Heterogomphus Burmeister, 1847 (Coleoptera: Scarabaeidae: Dynastinae: Oryctini) from Colombia”. Zootaxa 5590 (3). Auckland, New Zealand:345-64. https://doi.org/10.11646/zootaxa.5590.3.3.