Dehling, J. Maximilian. 2011. “A New Karst-Dwelling Species of Kalophrynus (Anura: Microhylidae) from Gunung Mulu National Park, Borneo, Malaysia”. Zootaxa 2737 (1). Auckland, New Zealand:49–60. https://doi.org/10.11646/zootaxa.2737.1.4.