Bravo-Valencia, L. and Rivera-Correa, M. (2011) “A new species of harlequin frog (Bufonidae: Atelopus) with an unusual behavior from Andes of Colombia”, Zootaxa. Auckland, New Zealand, 3045(1), pp. 57–67. doi: 10.11646/zootaxa.3045.1.4.