Coloma, L. A., Carvajal-Endara, S., Dueñas, J. F., Paredes-Recalde, A., Morales-Mite, M., Almeida-Reinoso, D., Tapia, E. E., Hutter, C. R., Toral, E. and Guayasamin, J. M. (2012) “Molecular phylogenetics of stream treefrogs of the Hyloscirtus larinopygion group (Anura: Hylidae), and description of two new species from Ecuador”, Zootaxa. Auckland, New Zealand, 3364(1), pp. 1–78. doi: 10.11646/zootaxa.3364.1.1.