Sathiandran, N. and Sabu, T. K. (2012) “New species, new synonym, and redescription of Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae) from the Western Ghats, India”, Zootaxa. Auckland, New Zealand, 3526(1), pp. 53–58. doi: 10.11646/zootaxa.3526.1.5.