Nunes, R. V. and Vaz-De-Mello, F. Z. (2016) “A new species of Holocephalus Hope from Paraguay (Coleoptera: Scarabaeidae: Scarabaeinae: Dichotomiini s. str.)”, Zootaxa. Auckland, New Zealand, 4136(2), pp. 397–400. doi: 10.11646/zootaxa.4136.2.12.