Cupello, M. and Vaz-De-Mello, F. Z. (2013) “Taxonomic revision of the South American dung beetle genus Gromphas Brullé, 1837 (Coleoptera: Scarabaeidae: Scarabaeinae: Phanaeini: Gromphadina)”, Zootaxa. Auckland, New Zealand, 3722(4), pp. 439–482. doi: 10.11646/zootaxa.3722.4.2.