Marinho, M. M. F., Carvalho, F. R., Langeani, F. and Tatsumi, F. L. (2008) “A new Hemigrammus Gill from upper rio Parana system, Southeastern Brazil (Characiformes: Characidae)”, Zootaxa. Auckland, New Zealand, 1724(1), pp. 52–60. doi: 10.11646/zootaxa.1724.1.2.