Rödel, M.-O., Schmitz, A., Pauwels, O. S. and Böhme, W. (2004) “Revision of the genus Werneria Poche, 1903, including the descriptions of two new species from Cameroon and Gabon (Amphibia: Anura: Bufonidae)”, Zootaxa. Auckland, New Zealand, 720(1), pp. 1–28. doi: 10.11646/zootaxa.720.1.1.