Génier, F. (2026) “Three new species of Onthophagus Latreille, 1802 from Bioko Island, Equatorial Guinea, belonging to d’Orbigny’s group IX, with a taxonomic note on O. denudatus d’Orbigny, 1902 (Coleoptera: Scarabaeidae: Scarabaeinae: Onthophagini)”, Zootaxa. Auckland, New Zealand, 5831(3), pp. 413–426. doi: 10.11646/zootaxa.5831.3.6.