Penz, C. M., Mohammadi, N. and Wahlberg, N. (2011) “Neotropical Blepolenis butterflies: wing pattern elements, phylogeny, and Pleistocene diversification (Lepidoptera, Nymphalidae)”, Zootaxa. Auckland, New Zealand, 2897(1), pp. 1–17. doi: 10.11646/zootaxa.2897.1.1.