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D. Vélez and F. Vivallo, “A new South American species of Centris (Heterocentris) Cockerell, 1899 with a key to the species with horn-like projections on the clypeus (Hymenoptera: Apidae: Centridini)”, Zootaxa, vol. 3357, no. 1, pp. 49–55, Jun. 2012.